Description

Key Characters:

Growth Form: Trees, shrubs, or sometimes lianas.

Stems:

Roots:

Leaves: Leaves simple. Opposite. Upper surfaces often with domatia; lower surfaces with pubescent domatia in axils of midrib and lateral veins. Pinnately veined. Stipules interpetiolar, leafy, distinct or adnate to petioles forming a sheath.

Flowers: Flowers in irregularly globose, ovoid to ellipsoid heads, bracts small. Flowers insect-pollinated, bisexual (perfect), heterostylous, or sometimes unisexual. Calyx lobes (2-)4-6 with open aestivation, lobes short, usually truncate, bases connate, rarely with 1-2 leaf-like lobes. Corolla funnelform or salverform, lobes (2-)4-6, valvate in bud, keeled within; nectary disk annular. Stamens as many as and alternate with the corolla lobes, inserted on corolla throat, exserted; anthers dithecal, opening by longitudinal slits, dorsifixed. Ovary inferior, occasionally partly so, or very rarely superior, 2-celled or incompletely 4-celled; ovules 1 per cell, attached below middle or near base; style terminal, slender, 2-branched, dimorphic, dry or occasionally wet.

Fruit: Drupes; those of the whole inflorescence connate to form a fleshy or hard syncarp; containing cartilaginous; 1-seeded pyrenes. Seeds 1 per fruit; obovoid or reniform; with well-developed oily endosperm; or endosperm occasionally scanty or absent.

Ploidy:

Habitat:

Elevation Range: